Central Taxonomic Thesaurus
Taxon Search
Taxon:
Display authors
Match on whole words
Display full tree below family
Display species with subgenera
  Animalia
      Arthropoda
        Hexapoda
               [Apterygota]
            Insecta
              Pterygota
                Neoptera
                       [Endopterygota]
                       [Exopterygota]
                    Coleoptera
                      Adephaga
                            Dytiscidae
                              Agabinae
                                     [Hydrotrupinae]
                                Agabini
                                     Agabus
                                       Agabus (Acatodes)
                                             Agabus ajax
                                             Agabus ancillus
                                             Agabus anthracinus
                                             Agabus approximatus
                                                   [Agabus prroximatus]
                                             Agabus arcticus
                                             Agabus audeni
                                             Agabus bicolor
                                             Agabus browni
                                             Agabus canadensis
                                             Agabus clypealis
                                             Agabus confinis
                                             Agabus congener
                                             Agabus coxalis
                                             Agabus discolor
                                             Agabus elongatus
                                                   [Agabus bryanti]
                                             Agabus fuscipennis
                                               Agabus fuscipennis fuscipennis
                                               Agabus fuscipennis ontarionis
                                                     [Agabus ontarionis]
                                             Agabus griseipennis
                                             Agabus hoppingi
                                             Agabus immaturus
                                             Agabus inexspectatus
                                             Agabus infuscatus
                                             Agabus inscriptus
                                             Agabus kootenai
                                             Agabus lutosus
                                             Agabus mackenziensis
                                             Agabus moestus
                                             Agabus morosus
                                             Agabus obliteratus
                                             Agabus obsoletus
                                               Agabus obsoletus nectris
                                                     [Agabus nectris]
                                               Agabus obsoletus obsoletus
                                             Agabus phaeopterus
                                             Agabus rumppi
                                             Agabus sasquatch
                                             Agabus smithi
                                             Agabus subfuscatus
                                             Agabus thomsoni
                                             Agabus zetterstedti
                                       Agabus (Agabus)
                                             [Carrhydrus]
                                             Agabus aeruginosus
                                             Agabus amnicola
                                                   [Agabus triton]
                                             Agabus antennatus
                                             Agabus bifarius
                                                   [Agabinus bifarius]
                                             Agabus clavicornis
                                                   [Agabus verus]
                                             Agabus colymbus
                                                   [Agabus coylmbus]
                                             Agabus crassipes
                                                   [Carrhydrus crassipes]
                                             Agabus disintegratus
                                                   [Agabus disintergratus]
                                             Agabus falli
                                                   [Agabus sharpi]
                                             Agabus gringo
                                             Agabus margaretae
                                             Agabus oblongulus
                                             Agabus pallens
                                                   [Agabus hudsonicus]
                                                   [Agabus zaitzevi]
                                             Agabus pisobius
                                             Agabus punctatus
                                             Agabus punctulatus
                                             Agabus taeniolatus
                                             Agabus undulatus
                                             Agabus xystrus
                                       Agabus (Gaurodytes)
                                             Agabus adpressus
                                                   [Agabus solus]
                                             Agabus affinis
                                             Agabus ambiguus
                                             Agabus austini
                                             Agabus erythropterus
                                             Agabus klamathensis
                                             Agabus leptapsis
                                             Agabus semipunctatus
                                             Agabus strigulosus
                                             Agabus tristis
                                             Agabus velox
                                             Agabus terminalis
                                     Agametrus
                                             Agametrus nitens
                                     Hydrotrupes
                                             Hydrotrupes palpalis
                                     Ilybiosoma
                                             Ilybiosoma amaroides
                                                   [Agabus amaroides]
                                             Ilybiosoma bjorkmanae
                                                   [Agabus bjorkmanae]
                                                   [Agabus rectus]
                                             Ilybiosoma brevicollis
                                                   [Agabus brevicollis]
                                             Ilybiosoma cordatum
                                                   [Agabus cordatus]
                                             Ilybiosoma flohrianus
                                                   [Agabus flohrianus]
                                             Ilybiosoma ilybiiformis
                                                   [Agabus ilybiiformis]
                                             Ilybiosoma lugens
                                                   [Agabus lugens]
                                             Ilybiosoma minnesotensis
                                                   [Agabus minnesotensis]
                                             Ilybiosoma oaxacaensis
                                                   [Agabus oaxacaensis]
                                             Ilybiosoma pandurus
                                                   [Agabus pandurus]
                                             Ilybiosoma perplexus
                                                   [Agabus perplexus]
                                             Ilybiosoma regularis
                                                   [Agabus regularis]
                                             Ilybiosoma roguus
                                                   [Agabus roguus]
                                             Ilybiosoma seriatum
                                                   [Agabus seriatus intersectus]
                                                   [Agabus seriatus seriatus]
                                                   [Agabus seriatus]
                                     Ilybius
                                             Ilybius angustior
                                             Ilybius ater
                                                   [Ilybius ungularis]
                                             Ilybius austrodiscors
                                             Ilybius biguttulus
                                                   [Ilybius laramaeus]
                                             Ilybius churchillensis
                                             Ilybius confertus
                                                   [Agabus confertus]
                                             Ilybius confusus
                                                   [Ilybius denikei]
                                             Ilybius discedens
                                             Ilybius discors
                                                   [Agabus discors]
                                             Ilybius erichsoni
                                                   [Agabus erichsoni]
                                             Ilybius euryomus
                                                   [Agabus euryomus]
                                             Ilybius fenestratus
                                             Ilybius fraterculus
                                             Ilybius fuliginosus
                                             Ilybius gagates
                                                   [Agabus gagates]
                                             Ilybius hypomelas
                                                   [Agabus hypomelas]
                                             Ilybius ignarus
                                             Ilybius incarinatus
                                             Ilybius jimzim
                                                   [Agabus jimzim]
                                             Ilybius larsoni
                                                   [Agabus larsoni]
                                             Ilybius lineellus
                                                   [Agabus lineellus]
                                             Ilybius oblitus
                                                   [Agabus oblitus]
                                             Ilybius opacus
                                                   [Agabus gelidus]
                                                   [Agabus opacus]
                                                   [Agabus pseudoconfertus]
                                             Ilybius picipes
                                             Ilybius pleuriticus
                                             Ilybius quadrimaculatus
                                             Ilybius subaeneus
                                             Ilybius vancouverensis
                                                   [Agabus vancouverensis]
                                             Ilybius vandykei
                                                   [Agabus vandykei]
                                             Ilybius verisimilis
                                                   [Agabus verisimillis]
                                             Ilybius vittiger
                                             Ilybius walsinghami
                                                   [Agabus walsinghami]
                                             Ilybius wasastjernae
                                                   [Agabus kenaiensis]
                                                   [Agabus palustris]
                                                   [Agabus wasastjernae]
                                     Leuronectes
                                             Leuronectes andinus
                                     Platambus
                                           [Agabinus]
                                             Platambus americanus
                                                   [Agabus americanus]
                                             Platambus apache
                                                   [Agabus apache]
                                             Platambus astrictovittatus
                                                   [Agabus astrictovittatus]
                                             Platambus aztec
                                                   [Agabus aztec]
                                             Platambus confusus
                                                   [Agabus confusus]
                                             Platambus flavovittatus
                                                   [Agabus flavovittatus]
                                             Platambus glabrellus
                                                   [Agabinus glabrellus]
                                             Platambus johannis
                                                   [Agabus johannis]
                                             Platambus maya
                                                   [Agabus maya]
                                             Platambus mexicanus
                                                   [Agabus mexicanus]
                                             Platambus obtusatus
                                                   [Agabus obtusatus]
                                             Platambus optatus
                                                   [Agabus optatus]
                                             Platambus planatus
                                                   [Agabus planatus]
                                             Platambus sculpturellus
                                                   [Agabinus sculpturellus]
                                             Platambus semivittatus
                                                   [Agabus semivittatus]
                                                   [Agabus texanus]
                                             Platambus spinipes
                                                   [Agabus spinipes]
                                             Platambus stagninus
                                                   [Agabus stagninus]
                                             Platambus texovittatus
                                                   [Agabus texovittatus]
                              Colymbetinae
                                Anisomeriini
                                     Anisomeria
                                             Anisomeria bistriata
                                Colymbetini
                                     Colymbetes
                                             Colymbetes dahuricus
                                                   [Colymbetes seminiger]
                                             Colymbetes densus
                                               Colymbetes densus densus
                                               Colymbetes densus inaequalis
                                                     [Colymbetes inaequalis]
                                             Colymbetes dolabratus
                                                   [Colymbetes dolobratus]
                                             Colymbetes exaratus
                                                   [Colymbetes rugipennis]
                                             Colymbetes fuscus
                                             Colymbetes incognitus
                                             Colymbetes paykulli
                                                   [Colymbetes longulus]
                                             Colymbetes sculptilis
                                             Colymbetes strigatus
                                     Hoperius
                                             Hoperius planatus
                                     Neoscutopterus
                                             Neoscutopterus angustus
                                             Neoscutopterus hornii
                                     Rhantus
                                       Rhantus (Nartus)
                                             Rhantus grapii
                                             Rhantus sinuatus
                                       Rhantus (Rhantus)
                                             Rhantus anisonychus
                                                   [Rhantus suffusus]
                                             Rhantus atricolor
                                             Rhantus binotatus
                                                   [Rhantus aequalis]
                                                   [Rhantus divisus]
                                                   [Rhantus flavogriseus]
                                             Rhantus bistriatus
                                             Rhantus calidus
                                             Rhantus consimilis
                                                   [Rhantus discedens]
                                                   [Rhantus tostus]
                                             Rhantus frontalis
                                                   [Rhantus notatus]
                                             Rhantus gutticollis
                                                   [Rhantus hoppingi]
                                                   [Rhantus hubbelli]
                                                   [Rhantus maculicollis]
                                             Rhantus oceanicus
                                             Rhantus pacificus
                                             Rhantus pseudopacificus
                                             Rhantus sericans
                                             Rhantus suturellus
                                                   [Agabus subopacus]
                                                   [Rhantus zimmermanni]
                                             Rhantus wallisi
                                                   [Ilybius wallisi]
                              Copelatinae
                                Copelatini
                                     Copelatus
                                       Copelatus (Copelatus)
                                             Copelatus anastomosans
                                             Copelatus biolleyi
                                             Copelatus blatchleyi
                                             Copelatus caelatipennis
                                               Copelatus caelatipennis angustatus
                                                     [Copelatus angustatus]
                                               Copelatus caelatipennis caelatipennis
                                               Copelatus caelatipennis fragilis
                                                     [Copelatus fragilis]
                                               Copelatus caelatipennis princeps
                                             Copelatus chevrolati
                                               Copelatus chevrolati chevrolati
                                               Copelatus chevrolati renovatus
                                             Copelatus debilis
                                             Copelatus distinctus
                                                   [Copelatus impressicollis]
                                             Copelatus glyphicus
                                                   [Copelatus decemstriatus]
                                                   [Copelatus punctulatus]
                                             Copelatus mancus
                                             Copelatus parvulus
                                                   [Copelatus mauiensis]
                                             Copelatus posticatus
                                     Lacconectus
                                             Lacconectus tonkinoides
                              Coptotiminae
                                Coptotimini
                                     Coptotomus
                                             Coptotomus difficilis
                                             Coptotomus interrogatus
                                                   [Coptotomus interrogatus interrogatus]
                                                   [Coptotomus interrogatus obscurus]
                                                   [Coptotomus obscurus]
                                             Coptotomus longulus
                                               Coptotomus longulus lenticus
                                                     [Coptotomus lenticus]
                                               Coptotomus longulus longulus
                                             Coptotomus loticus
                                             Coptotomus venustus
                              Dytiscinae
                                Acyliini
                                     Acilius
                                           [Acillius]
                                       Acilius (Acilius)
                                             Acilius abbreviatus
                                             Acilius athabascae
                                             Acilius fraternus
                                                   [Acilius fraternus dismalus]
                                                   [Acilius fraternus fraternus]
                                                   [Acillius fraternus]
                                             Acilius semisulcatus
                                                   [Acillius semisulcatus]
                                             Acilius simplex
                                             Acilius sylvanus
                                       Acilius (Homoelytrus)
                                             [Homaeolytrus]
                                             Acilius mediatus
                                                   [Acillius mediatus]
                                     Graphoderus
                                             Graphoderus cinereus
                                             Graphoderus fascicollis
                                                   [Graphoderus fasciatocollis]
                                             Graphoderus liberus
                                             Graphoderus manitobensis
                                             Graphoderus occidentalis
                                             Graphoderus perplexus
                                     Thermonectus
                                           [Thermonetus]
                                             Thermonectus basillaris
                                                   [Thermonectus basillaris intermedius]
                                                   [Thermonectus basillaris latecinctus]
                                                   [Thermonetus basillaris]
                                               Thermonectus basillaris basillaris
                                               Thermonectus basillaris galapagoensis
                                             Thermonectus circumscriptus
                                             Thermonectus marmoratus
                                                   [Thermonetus marmoratus]
                                             Thermonectus nigrofasciatus
                                               Thermonectus nigrofasciatus nigrofasciatus
                                               Thermonectus nigrofasciatus ornaticollis
                                                     [Thermonectus ornaticollis]
                                                     [Thermonetus ornaticollis]
                                             Thermonectus sibleyi
                                             Thermonectus succintus
                                                   [Thermonectus peninsularis]
                                Aubehydrini
                                       [Aubehydrinae]
                                     Notaticus
                                             Notaticus fasciatus
                                Cybistrini
                                     Cybister
                                       Cybister (Cybister)
                                             Cybister ellipticus
                                             Cybister explanatus
                                             Cybister fimbriolatus
                                               Cybister fimbriolatus crotchi
                                               Cybister fimbriolatus fimbriolatus
                                             Cybister flavocinctus
                                     Megadytes
                                       Megadytes (Bifurcitus)
                                             Megadytes giganteus
                                       Megadytes (Megadytes)
                                             Megadytes fraternus
                                Dytiscini
                                     Dytiscus
                                           [Leionotus]
                                           [Macrodytes]
                                             Dytiscus alaskanus
                                             Dytiscus carolinus
                                             Dytiscus circumcinctus
                                                   [Dytiscus anxius]
                                                   [Dytiscus ooligbucki]
                                             Dytiscus cordieri
                                             Dytiscus dauricus
                                                   [Dytiscus vexatus]
                                             Dytiscus fasciventris
                                             Dytiscus habilis
                                             Dytiscus harrisii
                                             Dytiscus hatchi
                                             Dytiscus hybridus
                                             Dytiscus marginalis
                                             Dytiscus marginicollis
                                             Dytiscus verticalis
                                Eretini
                                     Eretes
                                             Eretes sticticus
                                                   [Eretes occidentalis]
                                                   [Eretes stricticus]
                                Hydaticini
                                     Hydaticus
                                       Hydaticus (Guignotites)
                                             Hydaticus bimarginatus
                                       Hydaticus (Hydaticinus)
                                             Hydaticus xanthomelas
                                       Hydaticus (Hydaticus)
                                             Hydaticus aruspex
                                                   [Hydaticus modestus]
                                             Hydaticus cinctipennis
                                             Hydaticus piceus
                              Hydrodytinae
                                Hydrodytini
                                     Hydrodytes
                                             Hydrodytes dodgei
                              Hydroporinae
                                Bidessini
                                     Anodocheilus
                                             Anodocheilus exiguus
                                                   [Bidessus exiguus]
                                     Bidessonotus
                                             Bidessonotus browneanus
                                                   [Bidessus browneanus]
                                             Bidessonotus inconspicuus
                                                   [Bidessus inconspicuus]
                                             Bidessonotus longovalis
                                                   [Bidessus longovalis]
                                             Bidessonotus pulicarius
                                                   [Bidessus pulicarius]
                                                   [Bidessus subsericeus]
                                     Bidessus
                                             Bidessus unistriatus
                                     Brachyvatus
                                             Brachyvatus apicatus
                                                   [Bidessus seminulum]
                                                   [Brachyvatus seminulum]
                                     Comaldessus
                                             Comaldessus stygius
                                     Crinodessus
                                             Crinodessus amyae
                                     Liodessus
                                             Liodessus abjectus
                                                   [Liodessus youngi]
                                             Liodessus affinis
                                                   [Bidessus affinis affinis]
                                                   [Bidessus affinis]
                                                   [Uvarus affinis]
                                             Liodessus cantralli
                                             Liodessus crotchi
                                                   [Bidessus fuscatus]
                                                   [Hydroporus fuscatus]
                                                   [Liodessus fuscatus]
                                             Liodessus flavicollis
                                                   [Bidessus flavicollis]
                                                   [Uvarus flavicollis]
                                             Liodessus hobbsi
                                                   [Bidessus hobbsi]
                                             Liodessus noviaffinis
                                             Liodessus obscurellus
                                                   [Bidessus affinis erythrostomus]
                                                   [Bidessus affinis macularis]
                                                   [Bidessus affinis microreticulatus]
                                                   [Bidessus affinis nigrinus]
                                                   [Bidessus affinis obscurellus]
                                             Liodessus saratogae
                                     Neobidessus
                                             Neobidessus pulloides
                                             Neobidessus pullus
                                                   [Bidessus pullus]
                                               Neobidessus pullus floridanus
                                                     [Bidessus pullus floridanus]
                                               Neobidessus pullus pullus
                                                     [Bidessus pullus pullus]
                                             Neobidessus youngi
                                                   [Bidessus youngi]
                                     Neoclypeodytes
                                             Neoclypeodytes amybethae
                                             Neoclypeodytes cinctellus
                                                   [Bidessus cinctellus]
                                             Neoclypeodytes discretus
                                                   [Bidessus discretus]
                                             Neoclypeodytes fryii
                                                   [Bidessus decoratus]
                                                   [Neoclypeodytes decoratus]
                                             Neoclypeodytes leachi
                                                   [Bidessus leachi]
                                             Neoclypeodytes obesus
                                                   [Bidessus obesus]
                                             Neoclypeodytes ornatellus
                                                   [Bidessus ornatellus]
                                             Neoclypeodytes pictodes
                                                   [Bidessus pictodes]
                                             Neoclypeodytes plicipennis
                                                   [Bidessus plicipennis]
                                             Neoclypeodytes quadripustulatus
                                                   [Bidessus four-pustulatus]
                                                   [Bidessus quadripustulatus]
                                             Neoclypeodytes roughleyi
                                     Uvarus
                                             Uvarus amandus
                                                   [Bidessus amandus]
                                             Uvarus falli
                                                   [Bidessus falli]
                                             Uvarus granarius
                                                   [Bidessus granarius]
                                             Uvarus inflatus
                                                   [Bidessus inflatus]
                                             Uvarus lacustris
                                                   [Bidessus lacustris]
                                             Uvarus rogersi
                                                   [Bidessus rogersi]
                                             Uvarus spretus
                                             Uvarus subtilis
                                                   [Bidessus subplicatus]
                                                   [Bidessus subtilis]
                                             Uvarus suburbanus
                                                   [Bidessus suburbanus]
                                             Uvarus texanus
                                                   [Bidessus texanus]
                                Hydroporini
                                     Deronectes
                                             Deronectes aubei
                                  Deronectina
                                     Nectoboreus
                                             Nectoboreus aequinoctialis
                                                   [Boreonectes aequinoctialis]
                                     Haideoporus
                                             Haideoporus texanus
                                     Heterosternuta
                                           [Heterosternus]
                                             Heterosternuta alleghenianus
                                             Heterosternuta cocheconis
                                                   [Hydroporus cocheconis]
                                             Heterosternuta diversicornis
                                                   [Hydroporus diversicornis]
                                             Heterosternuta folkertsi
                                             Heterosternuta jeanneae
                                             Heterosternuta jenniferae
                                             Heterosternuta laetus
                                                   [Hydroporus laetus]
                                             Heterosternuta ohionis
                                             Heterosternuta oppositus
                                                   [Hydroporus oppositus]
                                             Heterosternuta ouachitus
                                             Heterosternuta phoebeae
                                             Heterosternuta pulcher
                                                   [Hydroporus pulcher]
                                             Heterosternuta sulphurius
                                                   [Hydroporus sulphurius]
                                             Heterosternuta wickhami
                                                   [Hydroporus wickhami]
                                     Hydrocolus
                                             Hydrocolus deflatus
                                                   [Hydroporus deflatus]
                                             Hydrocolus filiolus
                                                   [Hydroporus filiolus]
                                             Hydrocolus heggiensis
                                             Hydrocolus oblitoides
                                             Hydrocolus oblitus
                                                   [Hydroporus oblitus]
                                             Hydrocolus paugus
                                                   [Hydroporus paugus]
                                             Hydrocolus persimilis
                                                   [Hydroporus acadianus]
                                                   [Hydroporus aequus]
                                             Hydrocolus rubyae
                                             Hydrocolus rufiplanulus
                                                   [Hydroporus rufiplanulus]
                                             Hydrocolus rupinus
                                             Hydrocolus sahlbergi
                                             Hydrocolus stagnalis
                                                   [Hydroporus stagnalis]
                                     Hydroporus
                                             Hydroporus americanus
                                             Hydroporus appalachius
                                             Hydroporus aurora
                                             Hydroporus axillaris
                                             Hydroporus badiellus
                                             Hydroporus boreaorum
                                             Hydroporus brevicornis
                                             Hydroporus carri
                                             Hydroporus civicus
                                             Hydroporus columbianus
                                             Hydroporus contractulus
                                             Hydroporus dentellus
                                             Hydroporus despectus
                                             Hydroporus dichrous
                                                   [Hydroporus discicollis]
                                             Hydroporus falli
                                             Hydroporus fortis
                                             Hydroporus fuscipennis
                                             Hydroporus geniculatus
                                             Hydroporus glabriusculus
                                             Hydroporus gossei
                                             Hydroporus hirsutus
                                             Hydroporus hirtellus
                                             Hydroporus impressifrons
                                             Hydroporus klamathensis
                                             Hydroporus lapponum
                                                   [Hydroporus labradorensis]
                                             Hydroporus larsoni
                                             Hydroporus leechi
                                             Hydroporus longicornis
                                             Hydroporus longiusculus
                                             Hydroporus mannerheimi
                                                   [Hydroporus humeralis]
                                             Hydroporus melanocephalus
                                             Hydroporus melsheimeri
                                             Hydroporus morio
                                             Hydroporus nigellus
                                             Hydroporus niger
                                                   [Hydroporus latifrons]
                                             Hydroporus notabilis
                                                   [Hydroporus arcticus]
                                             Hydroporus obscurus
                                             Hydroporus occidentalis
                                             Hydroporus ophionis
                                             Hydroporus pervicinus
                                             Hydroporus planatus
                                             Hydroporus polaris
                                                   [Hydroporus subvirescens]
                                             Hydroporus puberulus
                                             Hydroporus rectus
                                             Hydroporus ruficeps
                                             Hydroporus rufilabris
                                                   [Hydroporus somnus]
                                             Hydroporus rufinasus
                                             Hydroporus sibiricus
                                             Hydroporus signatus
                                             Hydroporus similaris
                                             Hydroporus simplex
                                             Hydroporus sinuatipes
                                             Hydroporus spangleri
                                             Hydroporus striola
                                                   [Hydroporus striola striola]
                                                   [Hydroporus striola subtonsus]
                                             Hydroporus subpubescens
                                             Hydroporus tademus
                                             Hydroporus tartaricus
                                             Hydroporus tenebrosus
                                             Hydroporus transpunctatus
                                                   [Hydroporus funestus]
                                             Hydroporus tristis
                                             Hydroporus truncatus
                                             Hydroporus utahensis
                                             Hydroporus zackii
                                     Lioporeus
                                           [Falloporus]
                                             Lioporeus pilatei
                                                   [Hydroporus pilatei]
                                             Lioporeus triangularis
                                                   [Hydroporus triangularis]
                                     Nebrioporus
                                       Nebrioporus (Nebrioporus)
                                             [Potamonectes]
                                             Nebrioporus depressus
                                             Nebrioporus macronychus
                                             Nebrioporus rotundatus
                                     Neoporus
                                             Neoporus arizonicus
                                                   [Hydroporus arizonicus]
                                             Neoporus asyditus
                                             Neoporus aulicus
                                                   [Hydroporus aulicus]
                                             Neoporus baelus
                                             Neoporus blanchardi
                                                   [Hydroporus blanchardi]
                                             Neoporus carolinus
                                                   [Hydroporus carolinus]
                                                   [Hydroporus proximus]
                                             Neoporus cimicoides
                                                   [Hydroporus cimicoides]
                                             Neoporus clypealis
                                                   [Hydroporus clypealis]
                                             Neoporus dilatatus
                                                   [Hydroporus dilatatus]
                                             Neoporus dimidiatus
                                                   [Hydroporus dimidiatus]
                                                   [Hydroporus semirufus]
                                                   [Hydroporus solitarius]
                                             Neoporus dixianus
                                                   [Hydroporus dixianus]
                                             Neoporus effeminatus
                                                   [Hydroporus effeminatus]
                                             Neoporus floridanus
                                                   [Hydroporus floridanus]
                                             Neoporus gaudens
                                                   [Hydroporus gaudens]
                                             Neoporus hebes
                                                   [Hydroporus hebes]
                                             Neoporus helocrinus
                                             Neoporus hybridus
                                                   [Hydroporus hybridus]
                                             Neoporus latocavus
                                             Neoporus lecontei
                                                   [Hydroporus mixtus]
                                             Neoporus lobatus
                                                   [Hydroporus lobatus]
                                             Neoporus lynceus
                                                   [Hydroporus lynceus]
                                                   [Hydroporus republicanus]
                                             Neoporus mellitus
                                                   [Hydroporus mellitus]
                                             Neoporus pratus
                                             Neoporus psammodytes
                                             Neoporus rheocrinus
                                             Neoporus semiflavus
                                                   [Hydroporus semiflavus]
                                             Neoporus shermani
                                                   [Hydroporus shermani]
                                             Neoporus spurius
                                                   [Hydroporus spurius]
                                             Neoporus striatopunctatus
                                                   [Hydroporus striatopunctatus]
                                             Neoporus sulcipennis
                                                   [Hydroporus sulcipennis]
                                             Neoporus superioris
                                                   [Hydroporus superioris]
                                             Neoporus tennetum
                                             Neoporus tigrinus
                                                   [Hydroporus tigrinus]
                                             Neoporus undulatus
                                                   [Hydroporus consimilus]
                                                   [Hydroporus undulatus]
                                             Neoporus uniformis
                                                   [Hydroporus uniformis]
                                             Neoporus venustus
                                                   [Hydroporus venustus]
                                             Neoporus vitiosus
                                                   [Hydroporus vitiosus]
                                             Neoporus vittatipennis
                                                   [Hydroporus lineolatus]
                                                   [Hydroporus vittatipennis]
                                             Neoporus vittatus
                                                   [Hydroporus vittatus]
                                     Oreodytes
                                           [Deuteronectes]
                                             Oreodytes abbreviatus
                                                   [Deuteronectus abbreviatus]
                                                   [Hydroporus abbreviatus]
                                             Oreodytes alaskanus
                                                   [Hydroporus alaskanus]
                                                   [Hydroporus kincaidi]
                                                   [Hydroporus rainieri]
                                             Oreodytes alpinus
                                                   [Hydroporus alpinus]
                                             Oreodytes angustior
                                                   [Hydroporus angustior]
                                             Oreodytes congruus
                                             Oreodytes crassulus
                                                   [Hydroporus crassulus]
                                             Oreodytes humboldtensis
                                             Oreodytes laevis
                                                   [Hydroporus hortense]
                                                   [Hydroporus laevis]
                                                   [Hydroporus semiclarus]
                                                   [Hydroporus yukonensis]
                                                   [Oreodytes semiclarus]
                                             Oreodytes obesus
                                                   [Hydroporus obesus]
                                               Oreodytes obesus cordillerensis
                                               Oreodytes obesus obesus
                                             Oreodytes picturatus
                                                   [Deuteronectus picturatus]
                                                   [Hydroporus picturatus]
                                             Oreodytes productotruncatus
                                                   [Hydroporus productotruncatus]
                                             Oreodytes quadrimaculatus
                                                   [Hydroporus brodei]
                                                   [Hydroporus four-maculatus]
                                             Oreodytes recticollis
                                                   [Hydroporus recticollis]
                                             Oreodytes rhyacophilus
                                             Oreodytes sanmarkii
                                                   [Hydroporus rivalis]
                                             Oreodytes scitulus
                                               Oreodytes scitulus bisulcatus
                                                     [Hydroporus bisulcatus]
                                               Oreodytes scitulus scitulus
                                             Oreodytes septentrionalis
                                                   [Hydroporus septentrionalis]
                                             Oreodytes sierrae
                                             Oreodytes snoqualmie
                                                   [Hydroporus snoqualmie]
                                             Oreodytes subrotundus
                                                   [Hydroporus subrotundus]
                                     Paroster
                                             Paroster thapsinus
                                                   [Hygrotus thapsinus]
                                     Sanfilippodytes
                                             Sanfilippodytes adelardi
                                             Sanfilippodytes barbarae
                                                   [Hydroporus barbarae]
                                             Sanfilippodytes barbarensis
                                                   [Hydroporus barbarensis]
                                             Sanfilippodytes belfragei
                                                   [Hydroporus belfragei]
                                             Sanfilippodytes bertae
                                             Sanfilippodytes bidessoides
                                                   [Hydroporus bidessoides]
                                             Sanfilippodytes brumalis
                                             Sanfilippodytes compertus
                                             Sanfilippodytes corvallis
                                                   [Hydroporus corvallis]
                                             Sanfilippodytes edwardsi
                                                   [Hydroporus edwardsii]
                                             Sanfilippodytes hardyi
                                                   [Hydroporus hardyi]
                                             Sanfilippodytes kingii
                                                   [Hydroporus kingii]
                                             Sanfilippodytes latebrosus
                                                   [Hydroporus latebrosus]
                                             Sanfilippodytes malkini
                                                   [Hydroporus malkini]
                                             Sanfilippodytes pacificus
                                                   [Hydroporus pacificus]
                                             Sanfilippodytes palliatus
                                                   [Hydroporus palliatus]
                                             Sanfilippodytes planiusculus
                                                   [Hydroporus planiusculus]
                                             Sanfilippodytes pseudovilis
                                                   [Hydroporus pseudovilis]
                                             Sanfilippodytes rossi
                                                   [Hydroporus rossi]
                                             Sanfilippodytes sbordonii
                                             Sanfilippodytes setifer
                                             Sanfilippodytes terminalis
                                                   [Hydroporus browni]
                                                   [Hydroporus terminalis]
                                             Sanfilippodytes veronicae
                                             Sanfilippodytes vilis
                                                   [Hydroporus vilis]
                                             Sanfilippodytes williami
                                     Stictotarsus
                                             Stictotarsus aequinoctialis
                                                   [Hydroporus aequinoctialis]
                                                   [Hydroporus sericatus]
                                             Stictotarsus coelamboides
                                                   [Hydroporus coelamboides]
                                             Stictotarsus corvinus
                                                   [Hydroporus corvinus]
                                             Stictotarsus decemsignatus
                                                   [Hydroporus decemsignatus]
                                                   [Hydroporus mexicanus]
                                             Stictotarsus deceptus
                                                   [Deronectes deceptus]
                                                   [Hydroporus deceptus]
                                             Stictotarsus dolerosus
                                                   [Hydroporus dolerosus]
                                             Stictotarsus eximius
                                                   [Hydroporus addendus]
                                                   [Hydroporus eximius]
                                             Stictotarsus expositus
                                                   [Deronectes expositus]
                                                   [Hydroporus expositus]
                                             Stictotarsus falli
                                                   [Hydroporus corpulentus]
                                             Stictotarsus funereus
                                                   [Hydroporus funereus]
                                             Stictotarsus griseostriatus
                                                   [Deronectes griseostriatus]
                                                   [Hydroporus coloradensis]
                                                   [Hydroporus mathiasi]
                                                   [Potamonectes griseostriatus]
                                             Stictotarsus interjectus
                                             Stictotarsus minipi
                                             Stictotarsus neomexicanus
                                             Stictotarsus panaminti
                                                   [Deronectes panaminti]
                                                   [Hydroporus panaminti]
                                             Stictotarsus roffi
                                             Stictotarsus roffii
                                                   [Deronectes addendus]
                                                   [Deronectes roffi]
                                                   [Hydroporus pinguis]
                                                   [Hydroporus roffi]
                                             Stictotarsus spectabilis
                                             Stictotarsus spenceri
                                                   [Potamonectes spenceri]
                                             Stictotarsus striatellus
                                                   [Deronectes striatellus]
                                                   [Hydroporus striatellus]
                                             Stictotarsus titulus
                                                   [Hydroporus titulus]
                                     Suphrodytes
                                             Suphrodytes dorsalis
                                Hydrovatini
                                     Hydrovatus
                                             Hydrovatus brevipes
                                             Hydrovatus confertus
                                             Hydrovatus cuspidatus
                                             Hydrovatus davidis
                                             Hydrovatus hornii
                                                   [Hydrovatus horni]
                                                   [Hydrovatus major]
                                             Hydrovatus inexpectatus
                                             Hydrovatus peninsularis
                                             Hydrovatus platycornis
                                             Hydrovatus pustulatus
                                                   [Hydrovatus indianensis]
                                               Hydrovatus pustulatus compressus
                                               Hydrovatus pustulatus pustulatus
                                Hygrotini
                                     Hygrotus
                                       Hygrotus (Coelambus)
                                             Hygrotus artus
                                             Hygrotus berneri
                                             Hygrotus bruesi
                                             Hygrotus collatus
                                             Hygrotus compar
                                             Hygrotus curvipes
                                             Hygrotus dissimilis
                                                   [Hygrotus dispar]
                                             Hygrotus diversipes
                                             Hygrotus falli
                                             Hygrotus femoratus
                                             Hygrotus fontinalis
                                             Hygrotus fraternus
                                             Hygrotus fumatus
                                                   [Hygrotus sharpi]
                                             Hygrotus impressopunctatus
                                                   [Hygrotus impressopunctatus impressopunctatus]
                                                   [Hygrotus impressopunctatus lineellus]
                                             Hygrotus infuscatus
                                                   [Hygrotus dentiger]
                                                   [Hygrotus oregonus]
                                             Hygrotus laccophilinus
                                             Hygrotus lutescens
                                                   [Hygrotus medialis]
                                             Hygrotus marklini
                                                   [Hygrotus canadensis]
                                                   [Hygrotus virgo]
                                             Hygrotus masculinus
                                                   [Hygrotus fastidiosus]
                                             Hygrotus nigrescens
                                             Hygrotus novemlineatus
                                                   [Hygrotus hudsonicus]
                                             Hygrotus nubilus
                                                   [Hygrotus nubilis]
                                             Hygrotus obscureplagiatus
                                                   [Hygrotus columbianus]
                                             Hygrotus patruelis
                                                   [Hygrotus borealis]
                                                   [Hygrotus quebecensis]
                                             Hygrotus pedalis
                                             Hygrotus picatus
                                             Hygrotus punctilineatus
                                             Hygrotus salinarius
                                             Hygrotus sellatus
                                             Hygrotus semivittatus
                                             Hygrotus suturalis
                                                   [Hygrotus ovoideus]
                                             Hygrotus sylvanus
                                             Hygrotus thermarum
                                                   [Hygrotus bonnelli]
                                             Hygrotus tumidiventris
                                             Hygrotus turbidus
                                             Hygrotus unguicularis
                                             Hygrotus wardi
                                       Hygrotus (Hygrotus)
                                             Hygrotus acaroides
                                             Hygrotus farctus
                                             Hygrotus hydropicus
                                             Hygrotus intermedius
                                             Hygrotus marginipennis
                                             Hygrotus sayi
                                                   [Hydroporus punctatus]
                                Hyphydrini
                                     Desmopachria
                                             Desmopachria aspera
                                             Desmopachria cenchramis
                                             Desmopachria convexa
                                             Desmopachria defloccata
                                             Desmopachria dispersa
                                             Desmopachria grana
                                             Desmopachria latissima
                                             Desmopachria leechi
                                             Desmopachria mexicana
                                             Desmopachria mutchleri
                                             Desmopachria portmanni
                                             Desmopachria seminola
                                     Pachydrus
                                             Pachydrus brevis
                                             Pachydrus globosus
                                             Pachydrus obniger
                                             Pachydrus princeps
                                Laccornini
                                     Laccornis
                                           [Agaporus]
                                             Laccornis conoideus
                                             Laccornis deltoides
                                             Laccornis difformis
                                             Laccornis etnieri
                                             Laccornis latens
                                             Laccornis nemorosus
                                             Laccornis oblongus
                                             Laccornis pacificus
                                             Laccornis schusteri
                                Methlini
                                     Celina
                                             Celina angustata
                                             Celina contiger
                                             Celina grossula
                                             Celina hubbelli
                                             Celina imitatrix
                                             Celina occidentalis
                                             Celina palustris
                                             Celina picea
                                             Celina slossoni
                                Vatellini
                                     Derovatellus
                                       Derovatellus (Derovatellus)
                                             Derovatellus lentus
                                               Derovatellus lentus floridanus
                                               Derovatellus lentus lentus
                                     Macrovatellus
                                             Macrovatellus mexicanus
                              Laccophilinae
                                Agabetini
                                     Agabetes
                                             Agabetes acuductus
                                Laccophilini
                                     Laccodytes
                                             Laccodytes pumilio
                                     Laccophilus
                                             Laccophilus biguttatus
                                                   [Hydroporus biguttatus]
                                             Laccophilus fasciatus
                                               Laccophilus fasciatus fasciatus
                                               Laccophilus fasciatus rufus
                                               Laccophilus fasciatus terminalis
                                                     [Laccophilus terminalis]
                                             Laccophilus gentilis
                                             Laccophilus horni
                                             Laccophilus inagua
                                             Laccophilus maculosus
                                               Laccophilus maculosus decipiens
                                               Laccophilus maculosus maculosus
                                                     [Laccophilus decipiens]
                                               Laccophilus maculosus shermani
                                                     [Laccophilus shermani]
                                             Laccophilus mexicanus
                                               Laccophilus mexicanus atristernalis
                                                     [Laccophilus atristernalis]
                                               Laccophilus mexicanus mexicanus
                                             Laccophilus minutus
                                                   [Dytiscus dinutus]
                                                   [Dytiscus minutus]
                                             Laccophilus oscillator
                                             Laccophilus pictus
                                               Laccophilus pictus coccinelloides
                                                     [Laccophilus coccinelloides]
                                               Laccophilus pictus insignis
                                                     [Laccophilus insignis]
                                               Laccophilus pictus pictus
                                             Laccophilus proximus
                                             Laccophilus quadrilineatus
                                                   [Laccophilus four-lineatus]
                                               Laccophilus quadrilineatus quadrilineatus
                                             Laccophilus salvini
                                             Laccophilus schwarzi
                                             Laccophilus sonorensis
                                             Laccophilus undatus
                                             Laccophilus vacaensis
                                               Laccophilus vacaensis chihuahuae
                                               Laccophilus vacaensis vacaensis
                              Matinae
                                Matini
                                     Matus
                                             Matus bicarinatus
                                             Matus leechi
                                             Matus ovatus
                                             Matus relictus