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  Animalia
      Arthropoda
        Hexapoda
               [Apterygota]
            Insecta
              Pterygota
                Neoptera
                       [Endopterygota]
                       [Exopterygota]
                    Coleoptera
                      Polyphaga
                        Scarabeiformia
                          Scarabaeoidea
                            Geotrupidae
                                     Bolbocerastes
                                               Bolbocerastes imperialis imperialis
                                             Bolbocerastes regalis
                                             Bolbocerastes serratus
                              Bolboceratinae
                                     Bolbocerosoma
                                     Bolborhombus
                                             Bolborhombus sallaei
                                     Ceratotrupes
                                             Ceratotrupes bolivari
                                             Ceratotrupes fronticornis
                                     Eucanthus
                                             Eucanthus impressus
                                             Eucanthus lazarus
                                     Frickius
                                     Geotrupes
                                             Geotrupes balyi
                                               Geotrupes blackburnii blackburnii
                                               Geotrupes blackburnii excrementi
                                             Geotrupes opacus
                                             Geotrupes semiopacus
                                               Geotrupes splendidus miarophagus
                                               Geotrupes splendidus splendidus
                              Geotrupinae
                              Lethrinae
                                     Lethrus
                                     Megatrupes
                                             Megatrupes cavicollis
                                                   [Geotrupes cavicollis]
                                     Odonteus
                                             Odonteus obesus
                                     Taurocerastes