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  Animalia
      Arthropoda
        Hexapoda
               [Apterygota]
            Insecta
              Pterygota
                Neoptera
                       [Endopterygota]
                       [Exopterygota]
                    Coleoptera
                      Polyphaga
                        Scarabeiformia
                          Scarabaeoidea
                            Lucanidae
                              Aesalinae
                                     Apterocyclus
                                             Apterocyclus honoluluensis
                              Lampriminae
                              Lucaninae
                                     Lucanus
                                             Lucanus mazama
                              Nicaginae
                              Penichrolucaninae
                                     Pseudolucanus
                                             Pseudolucanus capreolus
                              Syndesinae