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  Animalia
      Arthropoda
        Chelicerata
            Arachnida
              Acari
                     [Acarina]
                  Acariformes
                    Sarcoptiformes
                      Oribatida
                             [Oribatei]
                        Brachypylina
                          Otocepheoidea
                            Tetracondylidae
                                     Afrotocepheus
                                       Afrotocepheus (Afrotocepheus)
                                             Afrotocepheus sinarmatus
                                       Afrotocepheus (Didierotocepheus)
                                             Afrotocepheus berndi
                                     Borneremaeus
                                             Borneremaeus hauseri
                                     Bulbocepheus
                                             Bulbocepheus hauserorum
                                     Cavernocepheus
                                             Cavernocepheus acutus
                                             Cavernocepheus furcatus
                                             Cavernocepheus fusifer
                                             Cavernocepheus monstruosus
                                             Cavernocepheus monticola
                                             Cavernocepheus obliquus
                                             Cavernocepheus undulatus
                                     Dolicheremaeus
                                             Dolicheremaeus absolon
                                             Dolicheremaeus africanus
                                             Dolicheremaeus almerodai
                                             Dolicheremaeus alticola
                                             Dolicheremaeus alveolatus
                                             Dolicheremaeus amazonicus
                                             Dolicheremaeus andulauensis
                                             Dolicheremaeus angustus
                                             Dolicheremaeus aokii
                                             Dolicheremaeus attenuatus
                                             Dolicheremaeus auritus
                                             Dolicheremaeus baloghi
                                             Dolicheremaeus bartkei
                                             Dolicheremaeus bengalensis
                                             Dolicheremaeus bifidus
                                             Dolicheremaeus bolivianus
                                             Dolicheremaeus brasilianus
                                             Dolicheremaeus bruneiensis
                                             Dolicheremaeus capillatus
                                               Dolicheremaeus capillatus capillatus
                                               Dolicheremaeus capillatus neonominatus
                                             Dolicheremaeus capreolatus
                                             Dolicheremaeus carinatus
                                             Dolicheremaeus ceylonicus
                                             Dolicheremaeus cicatricosus
                                             Dolicheremaeus claviger
                                             Dolicheremaeus conjunctus
                                             Dolicheremaeus coronarius
                                             Dolicheremaeus crispus
                                             Dolicheremaeus curvisetus
                                             Dolicheremaeus cuspidatus
                                             Dolicheremaeus damoeoides
                                             Dolicheremaeus densefoveolatus
                                             Dolicheremaeus distinctus
                                             Dolicheremaeus dorni
                                             Dolicheremaeus duplicarinatus
                                             Dolicheremaeus duplicatus
                                             Dolicheremaeus elisabethae
                                             Dolicheremaeus elongatus
                                             Dolicheremaeus euaensis
                                             Dolicheremaeus eusebioi
                                             Dolicheremaeus fijiensis
                                             Dolicheremaeus fujikawae
                                             Dolicheremaeus furcatus
                                             Dolicheremaeus furcillatus
                                             Dolicheremaeus furcula
                                             Dolicheremaeus geminus
                                             Dolicheremaeus georgii
                                             Dolicheremaeus giganticus
                                             Dolicheremaeus grafatus
                                             Dolicheremaeus granulatus
                                             Dolicheremaeus gyroplicatus
                                             Dolicheremaeus heterotrichus
                                             Dolicheremaeus himalayensis
                                             Dolicheremaeus hirsutus
                                             Dolicheremaeus inaequalis
                                             Dolicheremaeus indicus
                                             Dolicheremaeus infrequens
                                               Dolicheremaeus infrequens amamiensis
                                               Dolicheremaeus infrequens coreanus
                                               Dolicheremaeus infrequens hachijoensis
                                               Dolicheremaeus infrequens infrequens
                                               Dolicheremaeus infrequens taiwanus
                                             Dolicheremaeus inopinatus
                                             Dolicheremaeus keralensis
                                             Dolicheremaeus krantzi
                                             Dolicheremaeus kummeri
                                             Dolicheremaeus lineatus
                                             Dolicheremaeus lineolatus
                                             Dolicheremaeus longipilus
                                             Dolicheremaeus lucidus
                                             Dolicheremaeus luxtoni
                                             Dolicheremaeus machadoi
                                             Dolicheremaeus magnus
                                               Dolicheremaeus magnus ghanensis
                                               Dolicheremaeus magnus iteratus
                                               Dolicheremaeus magnus longiseta
                                               Dolicheremaeus magnus magnus
                                             Dolicheremaeus malangensis
                                             Dolicheremaeus manifera
                                             Dolicheremaeus mariehammerae
                                             Dolicheremaeus markusi
                                             Dolicheremaeus mauritii
                                             Dolicheremaeus minor
                                             Dolicheremaeus montanus
                                             Dolicheremaeus murphyi
                                             Dolicheremaeus mutabilis
                                             Dolicheremaeus nasalis
                                             Dolicheremaeus nepalensis
                                             Dolicheremaeus nimbus
                                             Dolicheremaeus obsessus
                                             Dolicheremaeus obtusisetus
                                             Dolicheremaeus oginoi
                                             Dolicheremaeus ohmensis
                                             Dolicheremaeus orientalis
                                             Dolicheremaeus ornatus
                                             Dolicheremaeus pannosus
                                             Dolicheremaeus papuensis
                                             Dolicheremaeus pectinatus
                                             Dolicheremaeus perisi
                                             Dolicheremaeus perreti
                                             Dolicheremaeus philippinensis
                                             Dolicheremaeus phyllotrichus
                                             Dolicheremaeus plurisetus
                                             Dolicheremaeus porcinolus
                                             Dolicheremaeus praeoccupatus
                                             Dolicheremaeus pseudofurcatus
                                             Dolicheremaeus punctatus
                                             Dolicheremaeus pustulatus
                                             Dolicheremaeus renukae
                                             Dolicheremaeus repetitus
                                             Dolicheremaeus robustus
                                             Dolicheremaeus rubripedes
                                             Dolicheremaeus russiae
                                             Dolicheremaeus sabahnus
                                             Dolicheremaeus samarensis
                                             Dolicheremaeus semicapillatus
                                             Dolicheremaeus semilunatus
                                             Dolicheremaeus seychellensis
                                             Dolicheremaeus siamensis
                                             Dolicheremaeus simplex
                                             Dolicheremaeus singaporensis
                                             Dolicheremaeus speciosus
                                             Dolicheremaeus subsimilis
                                             Dolicheremaeus sulcatus
                                             Dolicheremaeus sumatranus
                                             Dolicheremaeus szentivanyi
                                             Dolicheremaeus taidinchani
                                             Dolicheremaeus tamurai
                                             Dolicheremaeus tricornutus
                                             Dolicheremaeus trimucronatus
                                             Dolicheremaeus varilobatus
                                             Dolicheremaeus variolatus
                                             Dolicheremaeus vilhenarum
                                               Dolicheremaeus vilhenarum barbatula
                                               Dolicheremaeus vilhenarum vilhenarum
                                             Dolicheremaeus vitraeus
                                             Dolicheremaeus wallacei
                                             Dolicheremaeus wallworki
                                             Dolicheremaeus wangi
                                             Dolicheremaeus yoshii
                                             Dolicheremaeus yunnanensis
                                     Fernandocepheus
                                             Fernandocepheus franzi
                                     Fissicepheus
                                       Fissicepheus (Fissicepheus)
                                             Fissicepheus chinensis
                                             Fissicepheus clavatus
                                             Fissicepheus claviopsis
                                             Fissicepheus coronarius
                                               Fissicepheus coronarius coronarius
                                               Fissicepheus coronarius koreensis
                                             Fissicepheus curvisetosus
                                             Fissicepheus elegans
                                             Fissicepheus mitis
                                             Fissicepheus mitratus
                                             Fissicepheus nakanei
                                             Fissicepheus ornithorrhynchus
                                             Fissicepheus sitnikovae
                                             Fissicepheus steinmanni
                                             Fissicepheus subclavatus
                                             Fissicepheus vicinus
                                       Fissicepheus (Psammocepheus)
                                             Fissicepheus amabilis
                                             Fissicepheus haradai
                                     Flagellocepheus
                                             Flagellocepheus sagittatus
                                     Hydroecocepheus
                                             Hydroecocepheus romeroi
                                     Leptotocepheus
                                             Leptotocepheus flagellifer
                                             Leptotocepheus macromucronatus
                                             Leptotocepheus mahunkai
                                             Leptotocepheus orientalis
                                             Leptotocepheus trimucronatus
                                     Longocepheus
                                             Longocepheus australis
                                             Longocepheus globosus
                                             Longocepheus longus
                                             Longocepheus youngai
                                     Lophotocepheus
                                             Lophotocepheus simplex
                                     Neotrichocepheus
                                             Neotrichocepheus tongaensis
                                     Papillocepheus
                                             Papillocepheus areolatus
                                             Papillocepheus decoratus
                                             Papillocepheus decorus
                                             Papillocepheus deficiens
                                             Papillocepheus heterotrichus
                                             Papillocepheus neotropicus
                                             Papillocepheus reductus
                                             Papillocepheus tuberculatus
                                     Paradolicheremaeus
                                             Paradolicheremaeus tabulatus
                                     Pentacepheus
                                             Pentacepheus malindangensis
                                     Plenotocepheus
                                       Plenotocepheus (Neotocepheus)
                                             Plenotocepheus colliger
                                             Plenotocepheus longipilus
                                       Plenotocepheus (Plenotocepheus)
                                             Plenotocepheus africanus
                                             Plenotocepheus crinitus
                                             Plenotocepheus delicatissimus
                                             Plenotocepheus dentatus
                                             Plenotocepheus discrepans
                                             Plenotocepheus mollicoma
                                             Plenotocepheus undatus
                                             Plenotocepheus verrucosus
                                     Pseudotocepheus
                                       Pseudotocepheus (Constrictocepheus)
                                             Pseudotocepheus coarctatus
                                             Pseudotocepheus curtiseta
                                             Pseudotocepheus geminatus
                                             Pseudotocepheus gobletus
                                             Pseudotocepheus monteithi
                                             Pseudotocepheus orientalis
                                             Pseudotocepheus punctatus
                                             Pseudotocepheus tenuiseta
                                       Pseudotocepheus (Pseudotocepheus)
                                             Pseudotocepheus amonstruosus
                                             Pseudotocepheus andinus
                                             Pseudotocepheus aokii
                                             Pseudotocepheus asymmetricus
                                             Pseudotocepheus australis
                                             Pseudotocepheus bacilliger
                                             Pseudotocepheus clavigerus
                                             Pseudotocepheus contractus
                                             Pseudotocepheus curtipilus
                                             Pseudotocepheus dentatus
                                             Pseudotocepheus flagellatus
                                             Pseudotocepheus foveolatus
                                             Pseudotocepheus granulatus
                                             Pseudotocepheus hammerae
                                             Pseudotocepheus hauseri
                                             Pseudotocepheus lanceolatus
                                             Pseudotocepheus lienhardi
                                             Pseudotocepheus medius
                                             Pseudotocepheus neonominatus
                                             Pseudotocepheus parallelus
                                             Pseudotocepheus pauliani
                                             Pseudotocepheus pauliensis
                                             Pseudotocepheus promissus
                                             Pseudotocepheus pygmaeus
                                             Pseudotocepheus radiatus
                                             Pseudotocepheus setiger
                                             Pseudotocepheus sexdentatus
                                             Pseudotocepheus sexidimorphus
                                             Pseudotocepheus shauni
                                             Pseudotocepheus sturmi
                                             Pseudotocepheus szentivanyorum
                                             Pseudotocepheus tolanaro
                                             Pseudotocepheus vicarius
                                     Seboetocepheus
                                             Seboetocepheus coronatus
                                     Spinotocepheus
                                             Spinotocepheus foveolatus
                                             Spinotocepheus javensis
                                             Spinotocepheus nigromaculatus
                                             Spinotocepheus tjibodensis
                                     Trichocepheus
                                             Trichocepheus auriculus
                                             Trichocepheus lamellatus
                                     Trichocondyla
                                             Trichocondyla multisetosa
                                     Wallworkodes
                                             Wallworkodes carinatus
                                             Wallworkodes hyalinus